List of Cirsium species
The following species in the flowering plant genus Cirsium, the plume thistles, are accepted by Plants of the World Online.[1] A 2022 molecular study reassigned many species to other genera, but Cirsium remains a speciose genus.[2] Already as a result of the genetic study Ackerfield et al. 2020, Cirsium and Carduus were determined to be polyphyletic in their current circumscriptions, suggesting to either "consolidate all taxa into one large genus (Carduus or Cirsium)," or "recognize each major clade as a genus (Carduus, Cirsium, Eriolepis, Notobasis, Picnomon, Silybum, and Tyrimnus)".[3] Bureš et al. 2023 examined ploidy, genome length, morphological similarity and other lines of evidence, reaching a similar conclusion; whereas the former study referred to a broader range of genera by the term "Carduus–Cirsium group", the latter was published on the heels of the separation of Lophiolepis and Epitrachys from Cirsium, so the focused shifted to their surprising result of Carduus and Tirymnus being nested within Cirsium.[4] Since further genetic evidence is needed to resolve the intergeneric relationships of the Carduinae, this list will be restricted to species in the current genus Cirsium.
North American section
- Cirsium altissimum (L.) Spreng.[3]
- Cirsium anartiolepis Petr.[3]
- Cirsium andersonii Petr.[3]
- Cirsium andrewsii Jeps.[3]
- Cirsium arizonicum (A.Gray) Petr.[3]
- Cirsium brevifolium Nutt.[3]
- Cirsium brevistylum Cronquist[3]
- Cirsium canescens Nutt.[3]
- Cirsium carolinianum (Walter) Fernald & B.G.Schub.[3]
- Cirsium clavatum (M.E.Jones) Petr.[3]
- Cirsium coahuilense Ownbey & Pinkava[3]
- Cirsium conspicuum Sch.Bip.[3]
- Cirsium crassicaule Jeps.[3]
- Cirsium cymosum (Greene) J.T.Howell[3]
- Cirsium discolor (Muhl. ex Willd.) Spreng.[3]
- Cirsium douglasii DC.[3]
- Cirsium eatonii B.L.Rob.[3]
- Cirsium edule Nutt.[3]
- Cirsium ehrenbergii Sch.Bip.[3]
- Cirsium engelmannii Rydb.[3]
- Cirsium flodmanii Arthur[3]
- Cirsium fontinale Jeps.[3]
- Cirsium grahamii A.Gray[3]
- Cirsium hydrophilum Jeps.[3]
- Cirsium inamoenum (Greene) D.J.Keil[3]
- Cirsium jorullense Spreng.[3]
- Cirsium lappoides Sch.Bip.[3]
- Cirsium longistylum R.J.Moore & Frankton[3]
- Cirsium mexicanum DC.[3]
- Cirsium mohavense (Greene) Petr.[3]
- Cirsium nuttallii DC.[3]
- Cirsium ochrocentrum A.Gray[3]
- Cirsium parryi Petr.[3]
- Cirsium occidentale (Nutt.) Jeps.[3]
- Cirsium ownbeyi S.L.Welsh[3]
- Cirsium perplexans Petr.[3]
- Cirsium pitcheri Torr. & A.Gray[3]
- Cirsium pringlei Petr.[3]
- Cirsium pulcherrimum (Rydb.) K.Schum.[3]
- Cirsium pumilum Spreng.[3]
- Cirsium quercetorum (A.Gray) Jeps.[3]
- Cirsium remotifolium DC.[3]
- Cirsium rhaphilepis Petr.[3]
- Cirsium rhothophilum S.F.Blake[3]
- Cirsium rydbergii Petr.[3]
- Cirsium scariosum Nutt.[3]
- Cirsium subcoriaceum (Less.) Sch.Bip.[3]
- Cirsium texanum Buckley[3]
- Cirsium tracyi Petr.[3]
- Cirsium turneri Warnock[3]
- Cirsium undulatum Spreng.[3]
- Cirsium velatum (S.Watson) Petr.[3]
- Cirsium vinaceum Wooton & Standl.[3]
- Cirsium virginianum Michx.[3]
- Cirsium wheeleri Petr.[3]
- Cirsium wrightii A.Gray[3]
Section Cephalonoplos (Neck.) DC.
- Cirsium arvense (L.) Scop.[5][3]
Section Cirsium
Subsection Acaulia Petrak
Subsection Cirsium
Series Cirsium × Lophiolepis Bureš
- ×Cirsium vulgare (Savi) Ten. (pro sp.)[3][6]
Series Creticum
- Cirsium creticum (Lam.) d'Urv.[3]
- Cirsium tymphaeum Hausskn.[3]
Series Heterophylla Kharadze
- Cirsium dealbatum M.Bieb.[5]
- Cirsium erisithales Scop.[3]
- Cirsium heterophyllum (L.) Hill[5][3]
Series sine nomine
- Cirsium monspessulanum (L.) Hill.[3]
- Cirsium oleraceum (L.) Scop.[3]
- Cirsium palustre (L.) Scop.[3]
- Cirsium rivulare (Jacq.) All.[5]
Subsection Echenais (Cass.) Petrak (pro sect.)
- Cirsium alberti Regel & Schmalh.[5]
- Cirsium badakhschanicum Kharadze[5]
- Cirsium echinus (M.Bieb.) Hand.-Mazz.[5]
- Cirsium × glabrifolium (C.Winkl.) Petr.[5]
- Cirsium obvallatum M.Bieb.[3]
- Cirsium sairamense O.Fedtsch. & B.Fedtsch.[5]
- Cirsium semenowii Regel[5]
- Cirsium sieversii (Fisch. & C.A.Mey.) Petr.[5]
- Cirsium simplex C.A.Mey.[5]
Subsection Orthocentrum (Cass.) Kharadze
Series Cana Kharadze
incertae sedis
- Cirsium alatum (S.G.Gmel.) Bobrov[5][4]
- Cirsium albowianum Sommier & Levier[5]
- Cirsium elbrusense Sommier & Levier[5]
- Cirsium elodes M.Bieb.[5]
- Cirsium greimleri Bureš[8]
- Cirsium hygrophilum Boiss.[5]
- Cirsium libanoticum DC.[5]
- Cirsium oblongifolium K.Koch[5]
- Cirsium pseudopersonata Boiss. & Balansa[5]
- Cirsium pubigerum (Desf.) DC.[5][4]
- Cirsium rhabdotolepis Petr.[5]
- Cirsium subinerme Fisch. & C.A.Mey.[5][4]
- Cirsium svaneticum Sommier & Levier[5]
- Cirsium sychnosanthum Petr.[5]
- Cirsium uliginosum (M.Bieb.) Fisch.[5]
- Cirsium waldsteinii Rouy[5]
Subsection Sinocirsium Kitam.
Series Japonica Kitam.
- Cirsium japonicum DC.[3]
- Cirsium kamtschaticum Ledeb. ex DC.[3]
- Cirsium maackii Maxim.[5]
- Cirsium nipponicum Makino[3]
- Cirsium pectinellum A.Gray[5][3]
Series Schantarensia Kitam.
- Cirsium schantarense Trautv. & C.A.Mey.[5]
Subsection Spanioptilon (Less.) Kitam.
Series Asiatica Kharadze
Series Vlassoviana Kharadze
- Cirsium vlassovianum Fisch. ex DC.[5]
Section Pseudo-eriolepis (Nakai) Kitam.
- Cirsium pendulum Fisch. ex DC.[5]
incertae sedis
- Cirsium acantholepis Petr.
- Cirsium acrolepis (Petr.) G.B.Ownbey
- Cirsium × affine Tausch
- Cirsium aidzuense Nakai ex Kitam.
- Cirsium akimontanum Kadota
- Cirsium akimotoi Kadota & Mas.Saito
- Cirsium × alpestre Nägeli
- Cirsium alpicola Nakai
- Cirsium alpis-lunae Brilli-Catt. & L.Gubellini
- Cirsium alsophilum (Pollini) Soldano
- Cirsium × ambiguum All.
- Cirsium amplexifolium (Nakai) Kitam.
- Cirsium aomorense Nakai
- Cirsium apoense Nakai
- Cirsium appendiculatum Griseb.
- Cirsium arctorhaphis G.L.Nesom
- Cirsium argyracanthum DC.
- Cirsium arisanense Kitam.
- Cirsium × aschersonii Čelak.
- Cirsium ashinokuraense Kadota
- Cirsium ashiuense Shun.Yokoy. & T.Shimizu
- Cirsium × ausserdorferi Hausm. ex Treuinf.
- Cirsium austrohidakaense Kadota
- Cirsium ayasii H.Duman & Dirmenci
- Cirsium babanum Koidz.
- Cirsium barnebyi S.L.Welsh & Neese
- Cirsium bertolonii Spreng.
- Cirsium bicentenariale Rzed.
- Cirsium × bifrons Arv.-Touv.
- Cirsium × bipontinum F.W.Schultz
- Cirsium bitchuense Nakai
- Cirsium bocoynense G.L.Nesom
- Cirsium boninense Koidz.
- Cirsium borealinipponense Kitam.
- Cirsium × boulayi E.G.Camus
- Cirsium × bourgaeanum Willk.
- Cirsium brachycephalum Juratzka
- Cirsium bracteiferum C.Shih
- Cirsium × braun-blanquetianum Lawalrée
- Cirsium × breunium Goller & Huter
- Cirsium brevicaule A.Gray
- Cirsium × breviscapum Eichenf.
- Cirsium buchwaldii O.Hoffm.
- Cirsium buergeri Miq.
- Cirsium calcareum (M.E.Jones) Wooton & Standl.
- Cirsium candelabrum Griseb.
- Cirsium × candolleanum Nägeli
- Cirsium canoasense G.L.Nesom
- Cirsium carniolicum Scop.
- Cirsium × celakovskyanum K.Knaf
- Cirsium centaureae (Rydb.) K.Schum.
- Cirsium cernuum Lag.
- Cirsium charkeviczii Barkalov
- Cirsium × chatenieri Legrand ex H.J.Coste
- Cirsium chellyense R.J.Moore & Frankton
- Cirsium chikabumiense Kadota
- Cirsium chikushiense Koidz.
- Cirsium chilaishanense S.S.Ying
- Cirsium chinense Gardner & Champ.
- Cirsium chionanthum G.L.Nesom & García-Mor.
- Cirsium chokaiense Kitam.
- Cirsium chrysacanthum (Ball) Jahand.
- Cirsium cilicicum P.H.Davis & Parris
- Cirsium ciliolatum (L.F.Hend.) J.T.Howell
- Cirsium coloradense (Rydb.) Cockerell ex Daniels
- Cirsium confertissimum Nakai
- Cirsium congdonii R.J.Moore & Frankton
- Cirsium congestissimum Kitam.
- Cirsium × connexum Kitam.
- Cirsium consociatum S.F.Blake
- Cirsium × csepeliense Borbás
- Cirsium culebraense Ackerf.
- Cirsium dasyphyllum G.L.Nesom
- Cirsium dender Friis
- Cirsium dipsacolepis Matsum.
- Cirsium dissectum (L.) Hill
- Cirsium × dominii M.Schulze
- Cirsium domonii Kadota
- Cirsium drummondii Torr. & A.Gray
- Cirsium ducellieri Maire
- Cirsium durangense (Greenm.) G.B.Ownbey
- Cirsium dyris Jahand. & Maire
- Cirsium eliasianum Kit Tan & Sorger
- Cirsium englerianum O.Hoffm.
- Cirsium eorythros G.L.Nesom
- Cirsium epiroticum Petr.
- Cirsium eriophoroides (Hook.f.) Petr.
- Cirsium × erisithaliforme Preissm.
- Cirsium × erucagineum (Lam.) DC.
- Cirsium excelsius Petr.
- Cirsium × fabium Porta ex Kern.
- Cirsium falconeri (Hook.f.) Petr.
- Cirsium fanjingshanense C.Shih
- Cirsium fargesii (Franch.) Diels
- Cirsium faucium Petr.
- Cirsium fauriei Nakai
- Cirsium ferum Kitam.
- Cirsium filipendulum Lange
- Cirsium flavisquamatum Kitam.
- Cirsium foliosum DC.
- Cirsium forrestii (Diels) H.Lév.
- Cirsium × freyerianum W.D.J.Koch
- Cirsium funagataense Kadota
- Cirsium funkiae Ackerf.
- Cirsium furusei Kitam.
- Cirsium fuscotrichum C.C.Chang
- Cirsium gaditanum Talavera & Valdés
- Cirsium ganjuense Kitam.
- Cirsium glaberrimum (Petr.) Petr.
- Cirsium glabrum DC.
- Cirsium × grandiflorum Kitt.
- Cirsium grandirosuliferum Kadota
- Cirsium gratiosum Kitam.
- Cirsium grayanum Nakai
- Cirsium griffithii Boiss.
- Cirsium griseum (Rydb.) K.Schum.
- Cirsium × grossheimii Petr.
- Cirsium gyojanum Kitam.
- Cirsium hachijoense Nakai
- Cirsium hachimantaiense Kadota
- Cirsium hagurosanense Kadota
- Cirsium handelii Petr.
- Cirsium happoense Kadota
- Cirsium harrisonii (S.L.Welsh) Ackerf. & D.J.Keil
- Cirsium hasunumae Kadota
- Cirsium × heerianum Nägeli
- Cirsium heiianum Koidz.
- Cirsium helenioides (L.) Hill
- Cirsium henryi (Franch.) Diels
- Cirsium hesperium (Eastw.) Rydb.
- Cirsium heterotrichum Pančić
- Cirsium × heuseri Uechtr.
- Cirsium hida-paludosum Kadota & Nagase
- Cirsium hidakamontanum Kadota
- Cirsium homolepis Nakai
- Cirsium hookerianum Nutt.
- Cirsium horiianum Kadota
- Cirsium horridulum Michx.
- Cirsium hupingshanicum Z.C.Jin & Y.S.Chen
- Cirsium × hybridum W.D.J.Koch ex DC.
- Cirsium hygropodum G.L.Nesom
- Cirsium × iburiense Kitam.
- Cirsium imbricatum Petr.
- Cirsium inundatum Makino
- Cirsium ishizuchiense (Kitam.) Kadota
- Cirsium × ispolatovii Iljin ex Tzvelev
- Cirsium ito-kojianum Kadota
- Cirsium jaliscoense G.L.Nesom
- Cirsium joannae S.L.Welsh, N.D.Atwood & L.C.Higgins
- Cirsium kagamontanum Nakai
- Cirsium kasaianum Kadota
- Cirsium katoanum Kadota
- Cirsium kenji-horieanum Kadota
- Cirsium kirishimense Kadota & Mas.Saito
- Cirsium × kirschlegeri Sch.Bip.
- Cirsium kisoense (T.Yamaz. & K.Asano) Kadota
- Cirsium × kornhuberi Heimerl
- Cirsium × kozlovskyi Petr.
- Cirsium kubikialpicola Kadota
- Cirsium kujuense Kadota
- Cirsium latifolium Lowe
- Cirsium lecontei Torr. & A.Gray
- Cirsium leducii H.Lév.
- Cirsium leo Nakai & Kitag.
- Cirsium leucopsis DC.
- Cirsium lineare (Thunb.) Sch.Bip.
- Cirsium × linkianum M.Loehr
- Cirsium × lojkae Sommier & Levier
- Cirsium lomatolepis Petr.
- Cirsium longepedunculatum Kitam.
- Cirsium lucens Kitam.
- Cirsium luzoniense Merr.
- Cirsium magofukui Kitam.
- Cirsium mairei Halácsy
- Cirsium maritimum Makino
- Cirsium markaguntense (S.L.Welsh) Ackerf. & D.J.Keil
- Cirsium × martini L.C.Lamb.
- Cirsium maruyamanum Kitam.
- Cirsium masami-saitoanum Kadota
- Cirsium matsumurae Nakai
- Cirsium × mazanderanicum Petr.
- Cirsium × meratii P.Fourn.
- Cirsium michiliense G.L.Nesom
- Cirsium microspicatum Nakai
- Cirsium mimbresense G.L.Nesom
- Cirsium × mirabile Kitam.
- Cirsium × misawaense Nakai ex Kitam.
- Cirsium mollissimum G.L.Nesom
- Cirsium monocephalum H.Lév.
- Cirsium × moravicum Petr.
- Cirsium × muellneri Beck
- Cirsium muliense C.Shih
- Cirsium muramatsui Kadota
- Cirsium × murrianum Khek
- Cirsium muticum Michx.
- Cirsium myokoense Kadota
- Cirsium nagatoense Kadota
- Cirsium nagisoense Kadota
- Cirsium nambuense Nakai
- Cirsium nanhutashanense S.S.Ying
- Cirsium nasuense Kadota
- Cirsium neomexicanum A.Gray
- Cirsium × nevadense Willk.
- Cirsium nigriceps Standl. & Steyerm.
- Cirsium nippoense Kadota
- Cirsium nivale (Kunth) F.Dietr.
- Cirsium norikurense Nakai
- Cirsium × norrisii C.Bicknell
- Cirsium notorhaphis G.L.Nesom
- Cirsium novoleonense G.L.Nesom & García-Mor.
- Cirsium occidentalinipponense Kadota
- Cirsium × ochroleucum All.
- Cirsium ohminense Kadota
- Cirsium × okamotoi Kitam.
- Cirsium oligophyllum Matsum.
- Cirsium opacum (Kitam.) Kadota
- Cirsium orizabense Klatt
- Cirsium otayae Kitam.
- Cirsium pablillo G.L.Nesom
- Cirsium pascuarense (Kunth) Spreng.
- Cirsium × patens Kitam.
- Cirsium peckii L.F.Hend.
- Cirsium periacanthaceum C.Shih
- Cirsium × perplexissimum Kitam.
- Cirsium phulchokiense Kitam.
- Cirsium × pilosum Kitam.
- Cirsium pinetorum Greenm.
- Cirsium pitouchaoense S.S.Ying
- Cirsium × praealpinum Beck
- Cirsium praeteriens J.F.Macbr.
- Cirsium × prativagum Petr.
- Cirsium pseudocreticum (P.H.Davis & Parris) Yıldız, Dirmenci & Arabacı
- Cirsium pseudosuffultum Kadota
- Cirsium × pskemense Lazkov
- Cirsium pulchellum Wooton & Standl.
- Cirsium purpuratum Matsum.
- Cirsium × purpureum All.
- Cirsium pyrenaicum (Jacq.) All.
- Cirsium racemiforme Y.Ling & C.Shih
- Cirsium radians Benth.
- Cirsium rassulovii B.A.Sharipova
- Cirsium reglense Sch.Bip. ex Klatt
- Cirsium × reichardtii Jurotzka
- Cirsium × reichenbachianum M.Loehr
- Cirsium rengehydrophilum Kadota
- Cirsium repandum Michx.
- Cirsium rhinoceros Nakai
- Cirsium × rigens Wallr.
- Cirsium roseolum Gorl.
- Cirsium rosulatum Talavera & Valdés
- Cirsium rupinarum G.L.Nesom
- Cirsium × sabaudum M.Loehr
- Cirsium scapanolepis Petr.
- Cirsium schimperi (Vatke) C.Jeffrey
- Cirsium scopulorum (Greene) Cockerell
- Cirsium senjoense Kitam.
- Cirsium × sennenii Rouy
- Cirsium setidens (Dunn) Nakai
- Cirsium shansiense Petr.
- Cirsium shidokimontanum Kadota
- Cirsium shihianum Greuter
- Cirsium shimae Kadota
- Cirsium shinanense Shimizu
- Cirsium sichuanense Z.C.Jin & Y.S.Chen
- Cirsium sidi-guinii Pau & Font Quer
- Cirsium sieboldii Miq.
- Cirsium × siegertii Sch.Bip. ex Reichardt
- Cirsium × silesiacum Sch.Bip.
- Cirsium skutchii S.F.Blake
- Cirsium × soroksarense Wagner
- Cirsium souliei (Franch.) Mattf. ex Rehder & Kobuski
- Cirsium spicatum Matsum.
- Cirsium spinosissimum (L.) Scop.
- Cirsium spinosum Kitam.
- Cirsium spinuliferum (Kitam.) Kadota
- Cirsium stojanovii Kuzmanov
- Cirsium straminispinum C.Jeffrey
- Cirsium × subalpinum Gaudin
- Cirsium × subspinuligerum Peterm.
- Cirsium subuliforme G.B.Ownbey
- Cirsium × sudae E.Michálk. & Bureš
- Cirsium suffultum Matsum.
- Cirsium × sugimotoi Kitam.
- Cirsium × suspiciosum Beck
- Cirsium suzukaense Kitam.
- Cirsium suzukii Kitam.
- Cirsium swaticum Petr.
- Cirsium taiwanense Y.H.Tseng & Chih Y.Chang
- Cirsium takahashii Kadota
- Cirsium taliense (Jeffrey) H.Lév.
- Cirsium tamastoloniferum Kadota
- Cirsium tancitaroense G.L.Nesom
- Cirsium tanegashimense Kitam. ex Kadota
- Cirsium tashiroi Kitam.
- Cirsium tatakaense Y.H.Tseng, Y.H.Tseng, Chih Y.Chang & C.Y.Chang
- Cirsium × tataricum (L.) DC.
- Cirsium tenue Kitam.
- Cirsium tenuipedunculatum Kadota
- Cirsium tenuisquamatum Kitam.
- Cirsium tepehuanense G.L.Nesom
- Cirsium teshioense Kadota
- Cirsium tianmushanicum C.Shih
- Cirsium tioganum (Congdon) Petr.
- Cirsium togaense Kadota
- Cirsium tolucanum Petr.
- Cirsium townsendii (Petr.) G.L.Nesom
- Cirsium toyoshimae Koidz.
- Cirsium trachylomum S.F.Blake
- Cirsium tuberosum (L.) All.
- Cirsium uetsuense Kitam.
- Cirsium ugoense Nakai
- Cirsium umezawanum Kadota
- Cirsium unzenense Kadota & Mas.Saito
- Cirsium uzenense Kadota
- Cirsium valdespinulosum (Sennen) Sennen
- Cirsium valentinum Porta
- Cirsium vergelense G.L.Nesom
- Cirsium vernonioides C.Shih
- Cirsium verum Kadota
- Cirsium viperinum (D.J.Keil) Ackerf. & D.J.Keil
- Cirsium viridifolium (Hand.-Mazz.) C.Shih
- Cirsium × vivantii L.Villar, Segarra, J.López, Pérez-Coll. & Catalán
- Cirsium wakasugianum Kadota
- Cirsium wallichii DC.
- Cirsium × wankelii Reichardt
- Cirsium welwitschii Coss.
- Cirsium × wettsteinii Petr.
- Cirsium × wiedermannii Khek
- Cirsium × winklerianum Čelak.
- Cirsium × woodwardii (H.C.Watson) Nyman
- Cirsium × woronowii Petr.
- Cirsium yachiyotakashimae Kadota
- Cirsium yakusimense Masam.
- Cirsium yamauchii Kadota
- Cirsium yatsu-alpicola Kadota & Y.Amano
- Cirsium yezoalpinum H.Koidz. ex Kadota & S.Umezawa
- Cirsium yezoense (Maxim.) Makino
- Cirsium yoshidae Kadota
- Cirsium yuki-uenoanum Kadota
- Cirsium yuzawae Kadota
- Cirsium zamoranense Rzed.
- Cirsium zawoense Kadota
References
- "Cirsium Mill". Plants of the World Online. Royal Botanic Gardens, Kew. Retrieved 27 May 2023.
- Del Guacchio, Emanuele; Bureš, Petr; Iamonico, Duilio; Carucci, Francesca; De Luca, Daniele; Zedek, František; Caputo, Paolo (2022). "Towards a monophyletic classification of Cardueae: Restoration of the genus Lophiolepis (= Cirsium p.p.) and new circumscription of Epitrachys". Plant Biosystems. 156 (5): 1269–1290. Bibcode:2022PBios.156.1269D. doi:10.1080/11263504.2022.2131924. S2CID 252700781.
- Ackerfield, Jennifer; et al. (2020). "A prickly puzzle: Generic delimitations in the Carduus-Cirsium group (Compositae: Cardueae: Carduinae)". Taxon. 69 (4): 715–738. doi:10.1002/tax.12288.
- Bureš, Petr; et al. (2023-02-27). "Evolution of genome size and GC content in the tribe Carduinae (Asteraceae): rare descending dysploidy and polyploidy, limited environmental control and strong phylogenetic signal". Preslia. 95 (1): 185–213. doi:10.23855/preslia.2023.185. eISSN 2570-950X.
- Лукьяновна Харадзе, Анна (2001) [originally published 1963 in Russian]. "Cirsium". Flora Unionis Rerumpublicarum Socialisticarum Sovieticarum. Vol. 28. Translated by Dhote, A. K. Science publishers. pp. 51–213. ISBN 9781886106451.
- Bureš, Petr; et al. (2024-05-05). "Intergeneric hybrid origin of the invasive tetraploid Cirsium vulgare". Plant Biology. doi:10.1111/plb.13653. ISSN 1438-8677. PMID 38704835.
- Werner, Klaus (1976). "Cirsium Miller". Flora Europaea. Vol. 4. pp. 232–242. ISBN 0-521-08717-1.
- Bureš, Petr; Knoll, Aleš; Michálková, Ester; Šmarda, Petr; Šmerda, Jakub; Vavrinec, Martin (2018-05-09). "Cirsium greimleri: a new species of thistle endemic to the Eastern Alps and Dinarides". Preslia. 90 (2): 105–134. doi:10.23855/preslia.2018.105. eISSN 2570-950X.